Слайд 1 - Belarusian State University

Слайд 1 - Belarusian State University

6 1. 2. 3. 4. 5. 6. 7. 8. Reclinomonas mericana Elysia chlorotica

19 . 70- 20 Lynn Margulis Her theory that life evolved through cooperation and symbiosis, not through competition, was once scoffed at, and is now gaining widespread respect - ? -

( cob, Cox) -: Rhodospirillum, Bradyrhizobium, Rickettsia , Rickettsia prowazekii 2030 1 2-

Rickettsia prowazekii 1111523 .., 834 ; E.coli - 4288 24% . , . , , , . , - Jakobids Rick ettsi a pr o w az ekii

- icana r e m a o n as m o n i l

c Re

Jakobids - , R. mericana , 97 4 18 , R. mericana, . 4 rpo A-D. Jakoba libera rpo . str S12 S7 fusA tufA S10 E.coli R. americana

S12 S7 tufA S12 S7 M. polymorpha A. castellanii S10 L2 S19 S7 S3 L16 S10 L2 S19 S3 L16 S10

L2 S19 S3 L16 L2 S19 S3 L16 L3 L4 S12 S7 L23 spc E.coli R. americana

alpha L14 L24 L5 S14 S8 L6 L18 L14 L5 S14 S8 L6

L18 L5 S14 S8 L6 S13 S11 L5 S14 S8 L6 S13 S11 M. polymorpha A. castellanii

L29 S17 L14 S5 L30 L15 secY L36 S13 S11 secY S13 S11 S4 rpoA L17 rpoA R. mericana . 5 () 97 ( R. mericana). 1000 - (

1 ) AAAACTATAAGAAATAGTTATTTAAAAAAAAGGAAAAAAAAATGtufA TATGTGGGATTTCTTTGTGTGCTTAGAGAAAGGAATGTATATTTTATGnad1 TTAATGTTTTTAATATTTTTGATATAGTAAAGGAAAATAAGATG.nad11 TTAATTATTTAATATTTTTATTATTTTAAAAGGATTCTTATATGrps7 ATTCAAAGAAAATATTATAATAATTAATATGGGAATTTATTTTATG.rpoB ATAGAAGTGTTTGTATTTTTTAATATAAAAAGGTTGGTATAATATG nad4L GTTTCAAAACTTATTTTTAAGTAAAATGAAAGGATATTTATATG.cox2 TATAATTATAAATTCATAGAAAATAAAAAAAAGATAATCGCTTATG cox1 TATTGTATTTATAAATAATAAAAAAAACAAAGGAAATTCAAAATG..cox3 ****** 3 UUUCCUCAAGUAGGUCGA 16S Shine-Dalgarno R.americana. SD- 3- 16S R. mericana : 2- 12- , SD

! - Reclinomonas americana I II III IV V nad1, 2, 3, 4, 4L, 5, 6, 7, 8, 9, 10, 11 sdh 2, 3, 4 cob cox 1, 2, 3 atp 1, 6, 8, 9, 3 Core - -

/ Transporter SecY- ABC transporter rps 1, 2, 3, 4, 7, 8, 10, 11, 12, 13, 14, 19 rpl 2, 5, 6, 11, 14, 16, 1, 10, 18, 19, 20, 27, 31, 32, 34 tufA rpo A, B, C rpo D cox 11 secY yejR, U, V, W , , , , , . , Paramecium aurelium

3 - : , , mutS 90 - 13. II .

H+ H+ H+ ND1 ND2 ND3 ND6 ND4 ND5 ND4L H+ e- e- e- H2O O2 Cytb e-

COXI e- A8 A6 COXIII CoQ Cytc C: : : I II 7 III 0 >41 4

1 10 A A IV V 3 10 2 14 - cox2

. 2 , . : , . , Archaebacterium Archezoan () (+) A

- Archaebacterium () (+) B - , (rchezoan, ) - , - . () - (-) 80-

, : Archezoa Archeoprotista. , , . Archezoa , Giardia lamblia, , , , -

, Archeoprotista -, - , , : , : (, Entamoeba hystolitica) , 1 , , ??? ,

, , , , , Archezoa !

, - + ,

, , .. (A) , ( )

, , . -0,5 -1,5 -3,5 -3,8

, .. , , 2 , , . - 2 , . >1000.000.000.000 ,

, 1- , , , , 2700 6400 .

89 292 . Synechocystis 3229 200 - 50 Rhodophyta 59 33 Odontella (123) 18 77 Porphyra (182)

yanophora (128) 12 Euglena Marchantia 1 (57) 1 (83) Pinus Nicotiana (69) (76) Zea (76) Oryza (76) 5 (>205 )

1 110 4 3 7 1 (>2000 ) , 1-5% : Heterocapsa triquetra , , , , -, (2-10 ...). , ,

Heterocapsa triquetra - - - , 1- , 3 - 13 555 , 38 , - ?

( ); 3000 Synechocystis 46 24 , 44 ? : - - , ; - , , ( ); - - .

, , . . Placida cf. Dendritica , Elysia chlorotica , Vaucheria.

Pteraeolidia ianthina, , (zooxanthellae) - , 1-6 E. chlorotica

Vaucheria litorea (1-2 ) Vaucheria litorea Elysia chlorotica >8 ??? ??? : - ; Vaucheria ; , Vaucheria, ; Elysia ,

, , - , , Elysia . rbc , . . . n1 n2 C. , n2

() D. - n2 , , ?

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